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The existence of genetic correlations can be established by way of quantitative genetic breeding styles or via correlated phenotypic responses to selection. This suggests that lifespan and early reproduction are genetically negatively coupled, e.g. by way of antagonistic pleiotropic alleles (e.g., Flatt 2011, Fabian and Flatt 2011). But how can we reconcile the reality that VA is substantial whilst at the same time h2 is smaller? A most likely cause for the low heritability of life history traits is that, while VA (the numerator) is large, VP (the denominator) is much bigger than VA. A female North Pacific Giant Octopus (Enteroctopus dofleini) lives 3 Http://Www.Ftppro.Com/Gadgets2015/Rss_News.Php?A2=3467131045&A=Business+News&Id=46807&R=2&S=Http://Qooh.Me/Mrqrandell60876 to four years it lays thousands of eggs in a single bout and then dies. By contrast, a mature Coast Redwood Tree (Sequoia sempervirens) lives for numerous hundreds of years and produces millions of seeds every single year (Figure 1). As these two examples illustrate, organisms differ dramatically in how they develop, the time they take to develop, when they come to be mature, how lots of offspring of a unique size they make, and how long they reside. Collectively, the age-, size-, or stage-distinct patterns of development, growth, maturation, reproduction, survival, and lifespan define an organism's life cycle, its life history.

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